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It is powered by a new 5.2L V10 with direct injection, with improved power and torque. Unlike the precedent engine, this engine no longer has split crankpins, so it is not even firing, but Lamborghini says that due to the stiffer crankshaft it has less vibrations than the previous engine.

Transmission choices include six-speed manual or a six-speed E-gear, with E-gear revised to change gears by a rotational selector instead of a fore/aft movement, which offers 40% quicker shifts. E-gear also added Corsa setting and Thrust Mode launch control system.

The car was also 20 kg (44 lb) lighter than 2008 base Gallardo. Other changes include a new front bumper influenced by the Murcielago and exterior lights influenced by the Lamborghini Reventón.
Gallardo LP 560-4 has base price of US$201,000, but can go over $222,000 with e-gear, carbon ceramic brakes, navigation and other options.

Other factory options are available with Ad Personam program.

Lamborghini Gallardo